What age do cockerels loose fertility?

Cockerels being young male chickens generally have high fertility. I have had roosters remain fertile their entire life. I had a friend who had a closed flock of White Crested Black Polish that was derived from a single pair. These seriously inbred roosters tended to lose fertility around 3 or 4 years of age. Some heavy breed roosters retain fertility but become so arthritic that they can no longer effectively breed hens.
 

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