How am I getting feathered middle toes on offspring from parent birds with clean middle toes? If I understand the genetics properly, the Pti-1 is an allele that has 3 options: clean shanks, Langshan single toe feathering and Brahma two toe feathering. The Pti-1^B brahma type is supposed to be dominant. I have found this to be true with the Black Copper Marans. But with my Wheatens, I have paired up two parent birds with very clean middle toes - no stubs, nothing. And I'm getting 2 out of 3 offspring so far with feathered middle toes. I know there are shank feathering inhibitors that are recessive and exist in some production breeds that express clean shanks but possess the feathered shanks genes. But I have not read or heard of anything about an inhibitor that affects middle toe only. And it would have to be dominant to act this way. I'm stumped on this one. Any suggestions?