So they are going to do it again

Thanks, this is very helpful
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I went back and looked at the photos in post #18, which sparked much of this discussion. As nearly as I can tell, there is silvering on the wing feathers and the tail, but not as much on the feathers below the neck? Is that the area you are referring to as "wing saddles?"

What do you find to be the observable differences in birds who are carrying one WE gene as compared to two? Will just having one WE gene produce silvering on feathers (or dilution of color) or only a few white ocelli, with otherwise normal colored feathers?

Is the silvering coming from WE genes, or is there some other genetic mutation also turning up in birds who have WE genes?

Yes, that is the silver area. The degree can vary. WE birds have a frosted look to the wings. This frosted look can vary in degree. WE birds also can have white feathers in the wings and white feather tipping on the back. The degree of both varies with one WE gene or two WE genes.
 

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