- Nov 24, 2013
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I agree with the intended point of the post. 75% of a line when out crossing or 75% of a male/female when line breeding does not mean that 75% of the offspring’s genes came from that gene pool. I do think however, that more than a hundred years of selective breeding practices have proven that it greatly improves the odds that the random gene selection will come from those gene pools.
I'm having a hard time wrapping my head around the first statement; in my simple mind, it suggests that it is possible to hatch a giraffe, which I find highly unlikely. I suppose you mean 0 to 100% of the line they're 75% of.
I don’t think labels like 75 are essentially meaningless, just an ineffective way of describing the odds. Which can’t actually be described or calculated.
Actually there are ways to calculate the probabilities, as well as molecular genetic tools to discern exactly what percentage of an individual's genes came from a particular ancestor.
The problem is that past a certain point, genetic composition is more or less irrelevant to describe the complex type traits that are important to standard bred poultry. I would probably even argue that most of the discussion about 100 year old lines is past that point as well.
Of course the longer a population is selected toward the same goals, the more isogenic it becomes. But on the other hand, with selection based entirely on phenotype for complex genes, any and every gene and gene combination that results in that phenotype is selected, and that serves to preserve genetic diversity. Selection for vigor at the same time is also largely selection for heterozygosity, and that too slows the process of homogenizing the line.
But to make the 75% comment more clear, if a Ringlet line bird is crossed to another lines, all offspring contain 50% of their genes from each of the two lines. but when those F1s are backcrossed to the Ringlet line, the 50% of their genes that they contribute to their offspring is randomly sampled. On average they will contribute 50% of the genes from both parental lines, making the offspring 75/25, but this is only an average. They could contribute anywhere between 100/0 and 0/100, making the offspring anywhere between 100/0 and 50/50.
If two of those 50/50 "75"birds happen to be mated together, it is possible to create offspring with 0% Ringlet genes.
As I said before, the biggest limiter of the variability in these crosses is if the two lines were fairly genetically similar (to each other) to begin with. How standardized the two lines are (within themselves) is important, but not nearly as much.