Japanese quail dieing off

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Chirping
5 Years
My quail are gradually dieing off. They seem healthy then listless and then act paralyzed.

They are in 4'x2' pens with 3 birds per pen so I doubt overcrowding is the issue. Could this be fighting indunced? I think I have the agressive birds spread out but, it could be male on male mortality since the last three were all males.
 

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