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And that makes me think (sorry to go on) but maybe having ample headroom (like they would in the wild most times) would be better?

Duly, Happy Chooks, it's interesting that because mine stand/squat but maybe they would stand more upright if they had the headroom (the roof is low in the nestbox). Bok- bok- bkurkk after, which I've heard is an amouncement not of laying an egg but that timing is good for fertilisation (I.e...