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I'm fairly sure he hatched that way. Two or three of the Sussex I hatched has odd toes and I know it wasn't the incubation because I hatched five breeds in that same batch. I do believe it's a genetic thing-it's exactly like that Sussex boy I had, but he only had it on one toe.

His toes are not from perching on flat surfaces. Chickens don't grip with their rear toe like wild birds, so too round is actually not that great a perch. I hatched out one Speckled Sussex with one middle toe bent that way. Did you not notice it when he was smaller? I bet it's been that way and...