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As I understand it, estrogen plays a large role in it, as its presence naturally is high and causes fat in the liver. Chickens are obviously very hormonal. And then it just has something to do excess energy. Chickens who free range in general are less likely to get it regardless of diet...

Yes. I meal worm is equivalent to a snickers bar for us, which is why they love it. Overfeeding treats can lead to fatty livers and death in the birds.