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Most the time, yes. If only one could get them to wipe their feet before entering the nests. :gig Sand might help with that, or just track sand into the nests too.

What kind of bedding you use may depend on how you manage the manure. This is about cleaning, but covers my big picture -I use poop boards under roosts with thin(<1/2") layer of sand/PDZ mix, sifted daily(takes 5-10mins) into bucket going to friends compost. -Scrape big or wet poops off roost...