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This is from a web site. I can not load the reference. Tim

Quote: There may be 12 genes that cause brown egg color- brown eggs are genetically white also, so brown eggs are o+/o+. Brown egg pigments are normally added to the outside of the shell. Chickens that produce white eggs do not have any brown egg genes and are o+/o+, some have recessive sex...

Here is the chapter by Smyth on chicken genetics. This is from Crawfords book. Loaded with genetics of color. Download and save a copy. http://vision.berkeley.edu/wildsoet/Protocols/UsefulReferences/chicken_genetics-pp109-67.pdf Tim