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Adult cochins can't fly to save their souls. They are too heavy. Your smaller bodied birds,EE, PR can and will fly, 4' is easy for them, 5' takes some skill.

The main reason to cover the run is to keep overhead attacks out. If you have any birds of prey around, it's a good extra layer of security. I have young birds (brabanters and araucanas) that can easily fly 10 feet straight up. As they mature and get heavier they won't be able to but right...