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Actually, I wonder if a protein or other nutritional deficiency might be the problem. I have even read of cases where feather problems went away when the feed was changed.

Wow, hat's a mystery to me. The only thing that comes to mind is, do they have a wire bottom coop? Or maybe something else in their environment they could be rubbing on, or that could be that irritating to their skin. I suppose you could try some human antifungal cream, as for athlete's...