Does anyone know what the cause of this is? My quails joint is swollen and seems to have gotten worse, can barely stand up and tips.

I'd not let him suffer any longer. I'd off him so he doesn't suffer.
 
That is a great big enclosure for quail.. I’m surprised you don’t have more injured birds.. they really don’t need that much headroom or open length to run and take off, they have a big burst of momentum when they wanna really take off and fly the sudden stop or impacting another bird wide open will cause injury and death a lot of times..
 

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