Now this is where the emphasis on humidity gets overdone. Humidity is one contributing factor. Cold is a major factor.
Exposed surface area relative to volume of the comb, is also a major factor (that is, large plate-like single combs are much more vulnerable than small, thick pea combs).
I know of MANY cases of frostbite on single comb birds living in coops with excellent ventilation, sufficient space, and good litter management.
See the following thread, where someone actually measured humidity in the coop. This is in extremely cold climate, but I know of plenty of cases in my own much milder state of Mass.:
https://www.backyardchickens.com/t/722951/help-with-frostbite-alaska-coop
Yeah .. I shoulda worded that better, for sure: Clearly, there's a point where it's simply far too cold for chickens, no matter how dry it is. And, there are quite a few other factors, such as the specific reputations some breeds earn for cold hardiness, even beyond the aspects of their comb ... speaking of which, have never seen this worded better: "
Exposed surface area relative to volume of the comb ..."
But, for the very same reasons that you've so eloquently defined the variables of a specific chicken's comb, there's clearly just as direct a relationship between humidity and temperature within the lower range of temperatures as there is in the higher range (when heat stress becomes a threat). Air speed is also a factor, just as they calculate 'wind chill' when tellin' folks the weather. And, there'd be many other factors that'd have an affect.
But, that's what might help folks understand this better ... by studying the old-fashioned Temperature-Humidity Index, which has been in use for many years for predicting dangers from heat stress, the relationship can better be seen. And, I apologize for diggin' further, but I love mathematical equations, and developing ways to predict results, and studies that do this (which oughta be done for the lower side of a chicken's comfort zone ~'-)
From a study entitled:
Effect of Temperature-Humidity Index on Live Performance in Broiler Chickens*
*heat* stress related (edited to clarify ~'-)
The THI equations developed for poultry to date are shown in equations 1 through 4:
THIbroilers = 0.85 Tdb + 0.15 Twb (1, Tao and Xin, 2003)
THIlayers = 0.6 Tdb + 0.4 Twb (2, Zulovich and DeShazer, 1990)
THIhen turkeys = 0.74 Tdb + 0.26 Twb (3, Xin et al., 1992)
THItom turkeys = 0.42 Tdb + 0.58 Twb (4, Brown-Brandl et al., 1997)
where: THI = temperature-humidity index, °C
Tdb = dry-bulb temperature, °C
Twb = wet-bulb temperature, °C
I noticed that, in all but one of these equations? Tdb is given more weight than Twb, which may support your belief that temperature is of more importance. But, clearly, anything wet looses heat most quickly
:: edit :: when temperatures are low :: /edit ::, and would thereby freeze at higher temperatures. It'd sure be good to know what that range is, and for each specific breed ... speaking of which:
Henderson's Chicken Breed Chart is certainly worth careful consideration when choosing breeds, esp. where environments may be extreme.