On habitability:
I was able to keep 18 males and 24 females together in an 8ft x 8ft pen until 5 weeks, then I opened the connecting door to the other 1/2 where my existing 13 breeders, (2 roo/11 hen) and let them co-mingle. 55 quail in 128sqft. Everything was fine for a week then the hormones kicked in. At first I let it go, they have to get it out of their system before they can calm down. Then I saw 2 boys playing tug-of-war with a hen who was screaming a unique sound not normal to mating or being scared, they were really hurting her. I had to separate out all but 5 roos to calm things down. 15 boys in grow-out cages lost temporary mating rights until the cull, including one of my breeders.
Moral: If you have them together still and they are getting along, you are lucky and have a mellow group of boys, go with what works for you but always be prepared to intervene for the safety of all.
On laying:
I see that you have at least 2 factors keeping your numbers low, both have been identified.
1. Your hens are maturing at different rates. Last batch my first egg came at 5 weeks and 3 days, my latest-blooming hen took until just over 11 weeks to lay.
2. Lighting. 14-16 hours a day, enough said.
Again, stress of any kind can fudge your results even further, your roo/hen ratio may or may-not be a part of the equation.
I have barn owls attacking my aviary nightly. They can't get in but they try every night. Worst when they pace around on the galvanized roof. My birds are oblivious to them. It doesn't effect their laying at all. But when I muck out the enclosure they go on strike for a week.
Get in tune with your dynamics, everyone's are different. Environment, weather, temperament/personalities, predators. Some people can safely get away with things that the experts say are impossible.
Your flock dynamic as a whole can tell you what you are doing right and wrong if you know how to listen, or know what you are looking for.