Warning graphic who done it mystery

Predator must have came over the six foot welded wire fence.
I'd start by keeping predators out of the run. At least 75% of the ones you mentioned are climbers or can fly in. 6 feet means nothing to a climber. Ever watch a raccoon climb a tree? Cover the "roof" of the run with 1/2 inch hardware cloth if possible. Otherwise get electric fence and remove any trees or overhangs that a predator could climb and then drop down inside the run. Good luck!
 

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