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There was, possibly still is, a breed of chickens missing the long middle toe. The breed has no wings either because the genetics necessary for wings is believed to also be necessary for the middle toe. Peter Baumann of Des Moines visited the major chicken hatcheries and collected the wingless and nearly wingless culls of chicks and bred them for 12 years until he got wingless chickens which bred true. A dozen eggs were sent to England where they were hatched and used to create a flock of wingless birds with only 2 front toes. In 1924 W. L. Frank of Sherman, Texas developed a wingless chicken as did Dr. R.T. Renwald who developed a wingless strain in 1932.
In 1947 at the Iowa State Fair a strong wind would occasionally blow over the wingless chickens at The Register and Tribune's exhibit, and the gentleman operating the model train exhibit in the next booth was constantly standing up the wingless chickens as well as his model trains which got blown off the track.
Here is a link to a black and white video clip of the two-toed, wingless birds at the Shropshire Agricultural College in England in 1950.
http://www.britishpathe.com/video/wingless-chicks/query/chickens
Link to a pic of Peter Baumann looking over a flock of his wingless birds. Some of the birds clearly exhibit 3 toes.
http://www.gettyimages.com/detail/p...-baumann-high-res-stock-photography/tlp712099