As of today I haven't had any further issues.
I'm starting to wonder if they had a predator scare while I was at work one day.
It used to be that when I was opening up for the day before it was really light Rameses would stay up on the roost. Since the day of the issue, he's been flying down as soon as he hears me and refusing to let hens out into the run in the dark -- even chasing them back in if they went out.
This morning he jumped down off the roost when he heard me and then, when I went in so he was sure it was just me, went back to roost instead of patrolling the door.
I wonder if, in his limited chicken mind, the original crowding and even pecking behavior was his way to trying to get me to go into the safety of the coop instead of being out in the dangerous dark?
I had to turn my back on him a few times while working in the coop yesterday and he didn't do anything but continue to scratch in the straw I'd dropped while refilling the nests.
If the good behavior holds until the end of the month I'll let my husband and teens help with chores again.