Your intuition is correct, birds raised without a coop would not understand till they are taught to use a coop to sleep.No, they are not laying yet. I’d imagine sometime soon since they are now 4 months old. I also considered the ambient light being a contributing factor since a little bit does shine in the run. The roost bars are higher than the boxes and between the two I have 14 feet of space. I’ll have to find something to block them off this weekend. I was hoping they’d just correct it themselves but again they are new birds and I don’t believe they had a coop before.
You can use a bath towel or plastic sheeting if you still need to block the nest box. Just tack it on.