How do roosters learn to crow?

I believe it's instinct, with maybe a bit of mimicking thrown in if the opportunity is there. When my current cockerels first started crowing their cadence was exactly identical to my dominant rooster. As they perfected their crows they each learned to crow differently, so that now no two sound the same.
But in the past I've had single cockerels who learned to crow without ever hearing another rooster. Boys will all crow eventually no matter what :D
 

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