Curt: If the SP is homozygous (meaning both genes are SP/SP) then all the chicks will be SP, but they will carry a SD recessive (SP/SD). If a SP/SD is crossed to a SD/SD then half the chicks will be SP in color and half will be SD in color.
Everything I've read says the wild type e+ (the base of silver duckwing) is dominant to eb (the base of silver penciled / partridge) in the order of E locus dominance.
If that's the case, then wouldn't the F1 of a SP (eb/eb) x SDW (e+/e+) be SDW (presuming the duckwings are homozygous e+)?
If e+ dominantes eb, then they should phenotypically be e+ (SDW).
If the SDW is heterozygous (e+/eb), the cross of SDW (e+/eb) x SP (eb/eb) would give half heterozygous SDW (e+/eb), half SP (eb/eb).
I know Kari had it popping up and thought that was from Rex crossed to a carrier hen. Rex, iirc, is a birchen (ER), which would dominate e+ and would hide it. He (if he were ER/e+) would then produce heterozygous birchens (E/eb) and heterozygous SDW (e+/eb) when bred to a SP hen (eb/eb). He would create 3/4 Birchens and 1/4 SDW if bred to a hemizygous birchen carrying e+ (ER/e+). He should not be able to produce SP chicks, if that were the case, since both ER and e+ dominante eb. Since that's Kari's main cock bird for SPs of that generation which yielded a bunch of birchens and SPs with a small number of SDW, that makes it very unlikely, in my mind, that Rex carries e+. Makes me think her e+ snuck in with a birchen hen. Heterozygous birchen hen (ER/e+) x Rex (ER/eb) would yield birchens (ER/anything) and SDW chicks (e+/eb). That could make a batch of eggs from multiple hens show a bunch of birchen, SPs, and a few SD - these being the result of half the progeny of one hen.
Just thinking out loud, trying to reason it out...
Kari/Curt, can you help me out on what you've produced?
The tester of this would be has a SDW ever been produced from crossing two SP individuals with no chance a birchen snuck in there? Shouldn't be possible, if the order of dominance is as listed:
E, ER, ER- Fay, eWh, e+, eb, es, ebc, ey, eq
So we don't know yet what happens when a SP male covers a SDW female?
Presuming the above to be true,
SP = eb/eb S/S (ignoring the Pg for now)
SDW = e+/e+ S/-
Crossing these two should produce all e+/eb S/S males/ S/s- females.
e+/eb should be a duckwing appearance as e+ is listed as being more dominant than eb.
Crossing these F1s would yield 3/4 SDWs (1/3 e+/e+, 2/3 e+/eb) and 1/4 SPs (eb/eb).
If the SDWs are heterozygous, then the SP x SDW cross would yield 1/2 e+/eb (SDW) and 1/2 eb/eb (SP) in the F1.
I would bet, again, assuming the rule to be true, that Kari's SDW birds are heterozygous e+/eb birds. Breed the pair, and I'd expect 1/4 SP chicks in the offspring, with 3/4 SDW (2/3 of which carry SP).