Thanks for the infomation!@aart you give me too much credit. I certainly cannot fully explain but I'll try a bit. I could be wrong on part of this.
That spot on a yolk is the ovum (female sex cell with DNA) and enough nutrition to support the fertilized embryo until it can get hooked up right to absorb nutrition from the yolk and white. All eggs should have that. It takes a while for a fertilized embryo to send out blood vessels to start getting nutrition and do what it needs to to survive. So call it start-up kit in case the ovum is fertilized.
If the ovum gets fertilized by the sperm the egg still has about a day to go through the hen's internal egg making factory and it is around incubation temperature (technically a bit above). So the embryo is developing at a normal pace after fertilization but before it is laid and cools off. That ring around the blastoderm that makes it look like a bull's eye is the by-product of the use of nutrition while it is developing.
Probably more info than a lot of people need. I personally find it fascinating.