Silkie thread!

Okay, here is the down and dirty as I understand it thus far. Still working on fine tuning the phenotype/genotypes; for example, when there is red leakage does it indicate incomplete dominance of the silver gene (S/s) or an effect of autosomal red (AR/?) At this point, I cannot yet look at a bird and provide the genotype.

Anyway, feel free to correct:

First thing to remember, there are ONLY two pigments that make up color, Black (E-group) and Red (s and AR) White is the absence of pigment.

There are only five base colors in the black (the E group.) In order of dominance – E(black) = full extension of black, masks most patterning genes that may be present, ER (Birchen) = black with restriction i.e. allows patterning to show up including silver/gold leakage in hackle and saddle, fine lacing present in breast feathers, usually dark legs; eWh (Wheaten) = same as Duckwing (e) except no black lines in rooster hackle or saddle feathers and lighter down, secondary wing feathers remain black, females very different from (e) & (eb) with salmon of breast extending over all or most of body, chick down white or cream; Duckwing aka partridge or wild type (e) = salmon color breast on hens, black lines in hackle and saddle feathers of roosters; and last, Brown (eb) is same in appearance as (e) and (eWh) but will always have black shaft in hackle feathers, hens do not have the salmon breast of the wild type (e).

Red comes in a sex linked trait as gold (s) versus silver (S). Red also appears apart from silver/gold as autosomal red (Ar) which is dominant and not sex linked.

White comes in two masks – albino or “recessive” white masks the expression of ALL other colors and patterns which sometimes causes confusion because it acts dominant in this respect; however it is recessive because it only can express in a double dose (c/c). The “dominant” white (I) masks only the expression of black which is how you get white lacing instead of black or why you might want to breed to a black bird. You also only need one dose, hence the name dominant white.

Also at the locus for dominant white is the dun modifier (ID) which acts the same way as blue. When homozygous (ID/ID) it is khaki/dun, when heterozygous it is a chocolate color (ID/id) which is different than a true chocolate (choc), a homozygous recessive. Everything else is ‘not dun’ i.e. (id/id).

The rest of the alphabet soup affects the distribution of the two pigments (black and red) and sometimes the lack of all pigment (white), to various parts of the bird and the individual feathers. Some affect only black pigment like chocolate (choc), Melanotic (Ml) while others affect only red pigment like Mahogany (Mh) and dark brown (Db). Others yet affect all color pigments like Perlgray/Lavender (lav) which dilutes both pigments; the black to pale gray and red to straw gold or the sex linked barring (B) which turns expression of a pigment off and on over the length of a feather.

This is as much as I've got firm in my head. Still working out all the modifiers and how they interact. If you don't already have Sigrid's book, I have to say it is well worth the cost. And if you are breeding partridge and patterned birds, Grant Brereton's 21st Century Poultry Breeding is pretty good. I never knew you needed two breeding pens - 1 for exhibition male color and 1 for exhibition female color - until I read his book.
 
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Okay, here is the down and dirty as I understand it thus far. Still working on fine tuning the phenotype/genotypes; for example, when there is red leakage does it indicate incomplete dominance of the silver gene (S/s) or an effect of autosomal red (AR/?) At this point, I cannot yet look at a bird and provide the genotype.

Anyway, feel free to correct:

First thing to remember, there are ONLY two pigments that make up color, Black (E-group) and Red (s and AR) White is the absence of pigment.

There are only five base colors in the black (the E group.) In order of dominance – E(black) = full extension of black, masks most patterning genes that may be present, ER (Birchen) = black with restriction i.e. allows patterning to show up including silver/gold leakage in hackle and saddle, fine lacing present in breast feathers, usually dark legs; eWh (Wheaten) = same as Duckwing (e) except no black lines in rooster hackle or saddle feathers and lighter down, secondary wing feathers remain black, females very different from (e) & (eb) with salmon of breast extending over all or most of body, chick down white or cream; Duckwing aka partridge or wild type (e) = salmon color breast on hens, black lines in hackle and saddle feathers of roosters; and last, Brown (eb) is same in appearance as (e) and (eWh) but will always have black shaft in hackle feathers, hens do not have the salmon breast of the wild type (e).

Red comes in a sex linked trait as gold (s) versus silver (S). Red also appears apart from silver/gold as autosomal red (Ar) which is dominant and not sex linked.

White comes in two masks – albino or “recessive” white masks the expression of ALL other colors and patterns which sometimes causes confusion because it acts dominant in this respect; however it is recessive because it only can express in a double dose (c/c). The “dominant” white (I) masks only the expression of black which is how you get white lacing instead of black or why you might want to breed to a black bird. You also only need one dose, hence the name dominant white.

Also at the locus for dominant white is the dun modifier (ID) which acts the same way as blue. When homozygous (ID/ID) it is khaki/dun, when heterozygous it is a chocolate color (ID/id) which is different than a true chocolate (choc), a homozygous recessive. Everything else is ‘not dun’ i.e. (id/id).

The rest of the alphabet soup affects the distribution of the two pigments (black and red) and sometimes the lack of all pigment (white), to various parts of the bird and the individual feathers. Some affect only black pigment like chocolate (choc), Melanotic (Ml) while others affect only red pigment like Mahogany (Mh) and dark brown (Db). Others yet affect all color pigments like Perlgray/Lavender (lav) which dilutes both pigments; the black to pale gray and red to straw gold or the sex linked barring (B) which turns expression of a pigment off and on over the length of a feather.

This is as much as I've got firm in my head. Still working out all the modifiers and how they interact. If you don't already have Sigrid's book, I have to say it is well worth the cost. And if you are breeding partridge and patterned birds, Grant Brereton's 21st Century Poultry Breeding is pretty good. I never knew you needed two breeding pens - 1 for exhibition male color and 1 for exhibition female color - until I read his book.

Good raw base summary that still need some fine tuning.
It's not my place to make comments but maybe I can clear a thing or two.
-albino and recessive white are two different things. Albino have no pigment at all (example they have red eyes) . Recessive white have pigment (see your white Silkies having dark skin and eyes)
-recessive white .... "because it acts dominant". Here is intended recessive white act "epistatic" over other color-genes.

For the "all" modifiers you will need at least 100 years to work them all out !
1 example here is what modifiers are at work to make a Buff bird completely Buff without any trace of black ? We not know yet !
1 example for interaction is "single lacing" which is cause by the interaction of Co/Co (db+PgMl//db+PgMl) in linkage and crossing over inheritance.
If you are interested in this all I made a webpage on the basics (it's in Italian but contain 1000ths of photos that explain the text) :
https://sites.google.com/site/colorgenesandfeathers/
 
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Your questions may be leading me closer to an answer about a "sport" Ameraucana I hatched from someone else's stock. She hatched out of B/B/S eggs (blue daddy). She came out looking like the "pyle" pictured in Sigrid's Silkie book. Apparently, quite a few like her came out of that pen. I'm wondering if she is a leaky white. Now I've just re-read the section on dominant white in Sigrid's colour book. My hen also has the black/blue specks. When bred to a Blue Copper Marans cock, she threw what appeared to be a leaky (red) splash male bird. I'm wondering now if perhaps my Ameraucana female is carrying dominant white and that somewhere in her background a dominant white bird has been mislabeled as splash because they both appear to have Bl in them as well?

This is her ...


And this is her son ...
 
Your questions may be leading me closer to an answer about a "sport" Ameraucana I hatched from someone else's stock. She hatched out of B/B/S eggs (blue daddy). She came out looking like the "pyle" pictured in Sigrid's Silkie book. Apparently, quite a few like her came out of that pen. I'm wondering if she is a leaky white. Now I've just re-read the section on dominant white in Sigrid's colour book. My hen also has the black/blue specks. When bred to a Blue Copper Marans cock, she threw what appeared to be a leaky (red) splash male bird. I'm wondering now if perhaps my Ameraucana female is carrying dominant white and that somewhere in her background a dominant white bird has been mislabeled as splash because they both appear to have Bl in them as well?

This is her ...


And this is her son ...

The hen in the photo is in my opnion a e+/e+ s+/- Bl/Bl (a Splash partridge) look the "bluish" fearhers on the head and the salmon red chest but the phenotype of homozygous Blue => Splash (Bl/Bl) and heterozygous Domiant white (I/i+) can look very similar.
The cock in the other photo is (almost) sure a Splash but he is possible based on Silver (S/S or S/s+) with some autosmal red on his shoulder.

To be completely sure you can breed both to partridge.
When they are Bl/Bl X bl+/bl+ all chicks will be Bl/bl+ = Blue partridge
when they are I/i+ X i+/i+ 50% of the chicks will be partridge and 50% will be Pyle

What are B/B/S eggs ??
 
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Here are pictures of my silkie chicks I hatched out. All are from paint eggs from Twin Maple Ranch. I hatched out 16/18.

All of them
LL
Oh my goodness! A tub of happiness!! Soooo cute!
 
B/B/S = blue/ black/ splash
The hen in the photo is in my opnion a e+/e+ s+/- Bl/Bl (a Splash partridge) look the "bluish" fearhers on the head and the salmon red chest but the phenotype of homozygous Blue => Splash (Bl/Bl) and heterozygous Domiant white (I/i+) can look very similar.
The cock in the other photo is (almost) sure a Splash but he is possible based on Silver (S/S or S/s+) with some autosmal red on his shoulder.

To be completely sure you can breed both to partridge.
When they are Bl/Bl X bl+/bl+ all chicks will be Bl/bl+ = Blue partridge
when they are I/i+ X i+/i+ 50% of the chicks will be partridge and 50% will be Pyle

What are B/B/S eggs ??
 

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