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Interesting dilemma and responses. I have both breeds and will give you my 2 cents. Factor # 2 first as I'm understanding you will be selling purebred hatching eggs from this pen. Obviously you will be able to tell the Ameraucana eggs from the Marans eggs so it's fine to mix hens, but if you have an EE'er roo in the pen, you won't be able to know which eggs are pure. Regarding the popularity, as Henry has said, the Marans boom is over, however since they have been so badly bred to cash in on the boom, the few breeders who are maintaining good egg color and striving for correct conformation will be able to get decent prices for eggs. I believe the Ameraucanas are more the "rising stars" as people are becoming more educated as to true blue eggs and recognized varieties that breed true. IMO there is no chicken more beautiful than a W/BW roo or as lovely as the creamy beige hens.
1) The peacomb of the Ameraucana is less susceptible to frostbite.
3) Both my Marans and Ameraucanas started laying between 18 and 20 weeks. The Ameraucanas lay 5-6 eggs each a week, the Marans average 4. The Marans eggs are larger, though, and I've only had one broody...all the Ameraucanas have gone broody including one for the second time when her chicks were just 6 weeks old.
4 and 5) Both my Marans and Ameraucana roo's have shown no aggression towards people, and are good with their girls.When growing out in a pen mixed with other breeds, seem to establish dominance without too much brutality. I think either would be a good choice in this regard.
Another caveat is that my Marans are a little boring, while the Ameraucanas are friendlier and more fun to watch interact. It's like Junior High, the way some will be best bud's for awhile, then seem to be constantly having spats the next week. This may be my particular situation, as I've let the broodies raise babies in the same pen as the adults, and I guess the pecking order is constantly changing, but no one gets hurt and it's highly entertaining
. Even the adults though still fly up to roost on my shoulder when I'm in the pen. My Marans are much more wary.
My advice would be, since you are aiming for Olive Eggers, is to use the Marans in that pen to get the darker Olive color and pure Marans eggs. Find another spot for your EE'er and put that pretty Blue Wheaten roo in a separate pen with a W/BW girl or two to hatch some pure W/BW awesome chicks