Going in the nest boxes is actually a natural behaviour for roosters. They are to present the nest to the hen that he thinks is suitable for her to lay in, she inspects and then gives her yay or nay. My Blue Ameraucana roo use to do that with his favorit gals, he sometimes ended up sitting with them on the nest clucking away.
Yours is definitly a roo and if he is from a flock with multiple roos in it he probably was the bottom of the pecking order there for does not crow or breed the hens. Also if the hen is older and more dominant he may not breed her either cause she doesn't want to deal with a teenage boy practicing(got some cranky old bats like that myself, they only like their main guy and give youngsters the boot).
Also if you have an adult roo in your flock he may be again intimidated by him.
As for feathers, some patterns make feathers look pointy on some hens but on closer inspection they are round, if they are long, stringy, pointy with with those more see-through shiny edges they ARE rooster hackles.
Also bearded breeds of chickens don't get wattles like beardless breeds. In Ameraucana roosters they should downright be absent and not visible at all unless you dig through the beard.