Cute chicks.
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Would have loved to see that study.. however there is another way- it can be discovered by test-mating.
One easy way is to mate a NN to a non-naked necked bird. If the pair produces any non-naked chicks, it is proven heterozygote. All chicks having naked necks, probable homozygote- this proof becomes stronger the more chicks are hatched(10 at the very minimum.. 20-30 is far better). A good example of test-mating is breeding a showgirl to a silkie. If it produces both SG and silkies, then the showgirl is a heterozygote.
I have bred by now 300+ chicks from naked necked matings done various ways and I can assure that homozygotes can and do have either bowties or totally bare necks. In fact MOST homozygous naked necks have bowties, those would be the ones with small bowties with very visible naked areas below it and also exposed shoulders & crop. This visible difference is also apparent even in day olds- homozygotes have a pair of clearly separate and small puffs while heterozygote chicks have a much larger and "solid looking" fuzz patch on front of the necks. Once you recognize it, it becomes pretty easy to see.
There's always a "but". Some unrelated genes do affect the nakedness on a bird- peacomb is one. Peacomb also has the effect of reducing the overall number of feathers on a bird.. and also most heterozygous naked necks with peacombs look "very naked" exactly like the bird pictured above. I would dare guess that bird is a heterozygote despite being pretty naked, due to it also having a peacomb.
I have not done very many breedings with rosecombs, from what I see it appears rosecomb does make for more naked heterozygotes. It would explain why many heterozygote showgirls still have a fair amount of naked areas and also why homozyotes are bare necked, it's an added effect from the rosecomb. (most silkie combs are in actuality modified rosecombs).
Lastly, I tried breeding for total bare necks from(single combed) hatchery, backyard and various show stocks that had bowties, did not have success.. did manage to produce some birds that had only a single feather on each side of the neck but did not get any naturally bare necks.
Then in an egg trade, I got a batch of total backyard mongrels- leg feathering, crests, silky, etc. To my surprise those birds very easily produced total bare necks- even the single combed ones. Due to this group of birds I think there is something different about producing total bare necks(on single combed birds).. maybe there is another gene or ?? involved. They also prove the claim that only the "pure ones have totally bare necks" is false. Those birds were very obvious mutts.
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I have heard that claim several times too.. Often also I hear claim that the original stock had totally bare necks, however this bird shows a bowtie.