Fenrisulfr
Crowing
Roux is a dilute gene. Fee dilute is its own gene, and the gene itself is called Fee. They aren't related, and they're separate, so Roux doesn't cause Fee, and Fee doesn't cause Roux.
Fee is incompletely dominant so, like in that picture I posted of my two males, if a bird has even one copy, you can tell. I suppose it's possible that the Roux parents are also Fee. I'd have thought they'd be less red, though.
Here's some further reading on both Fee and Roux:
https://www.backyardchickens.com/th...ian-coturnix-quail-discussion-thread.1331206/
https://www.backyardchickens.com/threads/japanese-coturnix-falb-fee-discussion-thread.1331646/
I'm also wondering if the ones you're calling Roux are maybe not the same as what I'd call Egyptian - to me, a wild type bird with just Roux is Egyptian, which is this (borrowing a picture from Myshire):
Yours look a lot lighter, like they also have something else going on. Maybe they do have Fee which is why they're lighter. Would make sense, since if they're Fee, then instead of getting all Pharoah offspring, I'd expect you'd get all Falb Fee...which is what you got.
If you don't mind me borrowing the picture of your birds, I'll ask some quail genetics people what color they think they are.
Of course you can borrow the pics ...
... I bought the hatching eggs as "Roux Dilute".
I definately agree with you, that these have two factors. Roux is sex linked recessive and documentated. I didn't expected any Roux from that hatch. I just was expecting a dilution of the Pharaohs, but in another way as Fee ... but you see the result.
