The ones with contrasting edging are laced.
Penelope is partridge marked. With partridge coloring, the main feather color (here black) is on the outside of the feather, the contrasting color forms the next layer, then base then contrasting color again, then usually rest is base color. There are 2 variations on it if I remember correctly. I do not know what it is called when it isn't brown, though - but those are the same patterns as partridge marking.
@rural mouse knows or can find all RE: feather patterning/coloring